Douglas B. Walsh

Available Assistantships

PhD Assistantship for Integrated Pest Management on Hops

MS Assistantship in Alternative Leafhopper Control in Wine Grape Vineyards

MS Assistantship in Pollinator Protection and Enhancement (Alfalfa Seed)



Riparian Habitat/ Production Agriculture Interface:
I have several collaborative projects involving the study of arthropod population dynamics in disturbed and rehabilitated riparian habitats in the lower Yakima River watershed of south central Washington state. My major focus is developing the relationship among the host plants with constituent arthropod populations.

This research is being conducted in collaboration with Drs. Richard Zack (WSU Entomology) and Martin Williams (WSU Crop and Soil Science) and is supported by grants from the National Science Foundation Center for Integrated Pest Management, the Washington Tree Fruit Research Commission, and the Washington State Commission on Pesticide Registration.

Arthropod Pest Control on Washington Cropping Systems:
My laboratory is involved in multiple integrated pest management programs;

Crop / Pest(s) Cooperators Funding Agency
Apples / Lygus bug Mike Bush
(WSU Yakima County)
Washington Tree Fruit Research Commission
Stone Fruits / Lygus bugs,
western flower thrips
Mike Bush
(WSU Yakima County)
Washington Tree Fruit Research Commission
Peppermint, Spearmint / Spider Mites, Caterpillar Pests Washington State Mint Commission, Mint Industry Research Council, Washington State Commission on Pesticide Registration
Wine grapes / Cutworms Jack Watso
(WSU Benton County)
Wine Advisory Committee
Wine grapes / Mealy Bugs Ken Eastwell
(WSU Plant Pathology)
David James
(WSU Entomology)
Wine Advisory Committee & Washington State Commission on Pesticide Registration
Concord & other juice grapes /
Cutworms & Mealy Bugs
David James &
Jack Watson
Washington State Concord Grape Research Council
Dry bulb onions & onion seed / Western Flower Thrips Gary Pelter
(WSU Grant County)
Washington State Commission on Pesticide Registration, Columbia Basin Vegetable Seed Association, & Pacific Northwest Vegetable Association
Carrots/ Seed Corn Maggot Eric Sorenson
(WSU Franklin County)
Washington State Commission on Pesticide Registration


Extension Activities

Washington State IPM Coordinator:

  • Develop annual and 5 year plans of work for the implementation of IPM strategies in Washington State.
  • Participation in WCC-69, “Coordination of Integrated Pest Management Research and Extension/Outreach Programs for the Western States & Pacific Basin Territories”.
  • Manage IPM mini-grant program. Fiscal yr. 2000 for $69,000.

Washington State Liaison USDA IR-4 Project:

USDA/IR-4 is a federal/state/private cooperative that aspires to obtain clearances for pest control chemistries on minor crops. I serve as the State Liaison from Washington State to the to the USDA/ IR-4 program. My laboratory serves as the Field Research Center for EPA Region 10. EPA Region 10 consists of Eastern Oregon, Eastern Washington, and most of Idaho.

All IR-4 research–field and laboratory–is carried out according to Good Laboratory Practice (GLP) requirements as mandated by EPA. The research process is inspected by IR-4 quality assurance personnel to validate compliance.

Ex Officio Commissioner, Washington State Commission on Pesticide Registration:

Attend bi-monthly meetings throughout Washington State to review-approve/reject funding for pest management research projects.


  • Ph.D. – Univ. California at Davis, Entomology (1998)
  • B.S. – Univ. California at Santa Cruz, Biology, w/honors (1985)

Refereed Publications

*Nakawuka, P., R.T. Peters, S. Kenny, and D. Walsh.  2017.  Effect of deficit irrigation on yield quantity and quality, water productivity, and economic returns for four cultivars of hops in the Yakima Valley, Washington State.  Industrial Crops and Products. 98(2017) 82-92

*Boyle, N., S. Kesoju, S. Greene, & D. Walsh. 2016. Migratory beekeeping practices contribute insignificantly to transgenic pollen flow among fields of alfalfa produced for seed. Journal of Economic Entomology of four different cultivars of hops in the Yakima Valley, Washington State. J. Econ Etomol. doi: 10.1093/jee/tow243

Morales M. A., B. M. Mendoza, L. C. Lavine, M. D. Lavine, D. B. Walsh, F. Zhu 2016. Selection of Reference Genes for Expression Studies of Xenobiotic Adaptation in Tetranychus urticae. International Journal of Biological Sciences. 12(9): 1129-1139. doi: 10.7150/ijbs.16319

*Bahder, B.W., L.D. Bahder, M. Hauser, E. Beers & D.B. Walsh. 2016. Relative abundance and phenology of Drosophila Fallen, 1815 (Diptera: Drosophilidae) species in south-central Washington State. Pan-Pacific Entomologist, Vol. 92-2

Zhu, F., L. Lavine, S. O’Neal, C. Foss, & D. Walsh. 2016 Integrated Management of Insecticide Resistance in Urban Ecosystems. Insects 7, 2.

Walsh, D.B., S.D. O’Neal, A.E. George, D.P. Groenendale, R.E. Henderson, G.M. Groenendale, & M.J.

Hengel. 2016 Evaluation of Pesticide Residues from Conventional, Organic, and Non-treated Hops on Conventionally Hopped, Late-Hopped and Wet-Hopped Beers. Journal of the Association of Brewing Chemists.

*Piraneo, T. G., F. Zhu, J. Bull, M. D. Morales, L. C. Lavine, & D. B. Walsh. 2015 Molecular mechanisms of Tetranychus urticae chemical adaptation in hop fields. Sci. Rep. 5, 17090; doi:  10.1038/srep17090

*Bahder, B., L. D. Barrantes Barrantes, K. Hamby, M. Pascual F. Zalom, & D. B. Walsh. (2015). Population Genetics of Drosophila suzukii (Diptera: Drosophilidae) in Washington State and California. Environ. Entomol. DOI: 10.1093/ee/nvv117

Hull, J.J,  K. Chaney, S.M. Geib, J. A. Fabrick, C. S. Brent, D. Walsh, and L.Corley Lavine. 2014. Transcriptome-based identification of ABC transporters in the western tarnished plant bug Lygus Hesperus. PLoS 1. DOI: 10.1371/journal.pone.0113046

*Vinchesi, A.C. and D.B. Walsh. 2014. Quadrat Method for Assessing the Population Abundance of a Commercially Managed Native Soil-nesting Bee, Nomia melanderi (Hymenoptera: Halictidae) in Proximity to Alfalfa Seed Production in the Western USA”. J. Econ. Entom. 107: 1695-1699. DOI:

*Woods, J. L., A.J. Dreves, D.G  James, J.C Lee, D.B Walsh, &D.H. Gent. 2014. Development  Biological Control of Tetranychus urticae (Acari: Tetranychidae) and Phorodon humuli (Hemiptera: Aphididae) in Oregon Hop Yards. 107:570-581. DOI: 10.1603/EC13488

Peck, G. W., H. J. Ferguson, V. P. Jones, S. D. O’Neal, & D.B. Walsh. 2014. Use of a highly sensitive immunomarking system to characterize Face Fly (Diptera:Muscidae) dispersal from cow pats. Env. Entomol. 43: 116-122. DOI 10.1603/EN13139.

*Nakawuka, P., T.R. Peters, K.R. Gallardo, D. Toro-Gonzales, R. Okwany, & D.B. Walsh. 2014. Effects of deficit irrigation on yield, quality, and costs of production of native spearmint. J. Irrig Drain Eng. 140(5): 1-9. DOI:  10:1061/(ASCE)IR.1943-4774.0000719.

Steffan, S. A., J. C. Lee, M.E. Singleton, A. Vilaire, D. B. Walsh, Laura S. Lavine, & K. Patten. 2013. Susceptibility of cranberries to Drosophila suzukii  (Diptera: Drosophilidae). J. Econ Entomol.106:2424-2427. DOI:10.1603/EC13331

Lee, J.C.,  P. W. Shearer, L. D. Barrantes, E. H. Beers, H. J. Burrack, D. T. Dalton. A. J. Dreves, L, J. Gut, K. A. Hamby, D. R. Haviland, R. Isaacs, A. L. Nielsen, T. Richardson, C. R. Rodriguez-Saona,  C. A. Stanley, D. B. Walsh, V. M. Walton, W. L. Yee, F. G. Zalom, and D. J. Bruck. 2013. Improving trap design for monitoring Drosophila suzukii (Diptera: Drosophilidae) . Environ. Entomol.

Wu, M., H. Gotoh, T. Waters, D. B Walsh, L. C. Lavine. 2013 Identification of an alternative Knockdown Resistance kdr-like mutation, M918L, and a novel mutation, V1010A, in the Thrips tabaci voltage-gated sodium channel gene. Pest Management Sci. DOI: 10.1002/ps.3638

*Bahder, B.W., S. Poojari, O. J. Alabi, R. A. Naidu, & D. B. Walsh. 2013. Pseudococcus maritimus   (Hemiptera: Pseudococcidae) and Parthenolecanium corni (Hemiptera: Coccidae) are capable of transmitting Grapevine leafroll-associated virus 3 between Vitis labruscana and Vitis vinifera. Env. Entomol.

Peck, G. W., H. J. Ferguson, J.T. LePage, V. R. Hebert, S. D. O’Neal, & D.B. Walsh. 2013 Evaluation of Sunlight-Exposed Pyrethroid-Treated Netting for Control of Face Fly and House Fly (Diptera: Muscidae). Pest Man. Sci. DOI: 10.1002/ps.3537

*Bahder, B.W., O. Alabi, S. Poojari, D. B. Walsh, R. A. Naidu, 2013 A Survey for Grapevine Viruses in Washington State ‘Concord’ (Vitis x labruscana L.) Vineyards. Plant Health Prog. doi:10.1094/PHP-2013-0805-01-RS

*Buttar, B.S. , M L. Nelson, J. R. Busboom, D. D. Hancock, D. B. Walsh, D. P. Jasmere, 2013. Effect of heat treatment on viability of Taenia hydatigena eggs. Experimental Parasitology. 133:421–426

*Bahder, B. W. R. A. Naidu, K. M. Daane, J. G. Millar, &D. B. Walsh. 2013. Pheromone-based Monitoring of Pseudococcus maritimus (Hemiptera:Pseudococcidae) Populations in Concord Grape Vineyards. J. Econ,.Entomol. 106: 482-490

*Vinchesi, A., L. Lavine, D. Cobos, & D. Walsh. 2012 Manipulation of Soil Temperatures to Influence Brood Emergence in the Alkali Bee (Nomia melanderi). Apidologie. 10.1007/s13592-012-0180-7

*Zhou C, Kandemir I, Walsh D.B, Zalom F.G, Lavine L.C. 2012. Identification of Lygus hesperus by DNA Barcoding Reveals Insignificant Levels of Genetic Structure among Distant and Habitat Diverse Populations. PLoS ONE 7(3): e34528. doi:10.1371/journal.pone.0034528

Lee, J. C., H. J. Burrack, L. D. Barrantes, E.H. Beers, A. J. Dreves, K. A. Hamby,  D. R. Haviland, R. Isaacs, T. A. Richardson, P. W. Shearer, C. A. Stanley, D. B. Walsh, V. M. Walton, F. G. Zalom, and D. J. Bruck. 2012 Evaluation of monitoring traps for Drosophila suzukii (Diptera: Drosophilidae) in North America. J. Econ. Entomol. 105(4): 1350Ð1357

  1. M. Daane, M. C. Middleton, R. Sforza, M. L. Cooper, V. M. Walton, D.B. Walsh, T. Zaviezo, and R. P. P. Almeida. 2011 Development of a multiplex PCR for identification of vineyard mealybugs. Env. Entomol. 40:1595-1603

Dalton D.T., Walton V.M., Shearer, P.W., Walsh, D.B., Caprile J. and R. Isaacs. 2011. Laboratory survival of Drosophila suzukii under simulated winter conditions of the Pacific Northwest and seasonal field trapping in five primary regions of small and stone fruit production in the United States. Pest Manag Sci DOI 10.1002/ps.2280. 67: 1368 – 1374

*Okwany, R.O., and R.T. Peters. K. Ringer, D. Walsh, and M. Rubio.  2011.  Impact of Sustained Deficit Irrigation on Spearmint (Mentha spicata L.) Water Usage and Yield. Irrigation Science.

Walsh, D. B., M.P. Bolda, R. E. Goodhue, A.  J. Dreves, J.  Lee, D. J. Bruck, V. M. Walton, S. D. O’Neal, & F. G. Zalom. 2011. Drosophila suzukii (Diptera: Drosophilidae): Invasive Pest of Ripening Soft Fruit Expanding Its Geographic Range and Damage Potential. J. IPM 2:

*Waters, T.D., Zack, R.S. Zack, & D.B. Walsh. 2009. Creek Proximity and Flora Effects On Arthropod Populations in Riparian Buffers in Washington State, USA. Journal of Environmental Science and  Engineering. 4:52-60

*Weekes, L., Walsh, D., Ferguson, H.J., & Ross, C.  2009, Impact of multicolored Asian lady beetles on the sensory properties of Concord and Niagara grape juice.  Journal of Food Science : An Official Publication of the Institute of Food Technologists.  Dec 8. 2009

*Metzger, C., R. Boydston, H. Ferguson, M.M. Williams II, R. Zack, & D.Walsh. 2008. Interactions between population density of the Colorado potato beetle, Leptinotarsa decemlineata, and herbicide rate for suppression of solanaceous weeds. J. of Insect Sci. 8:38.

Walsh, D. & T.D. Waters. Thrips Infestations Increase Storage Rots in Dry Bulb Onions. 2008   Acta Hort.Proceedings of the World Edible Allium Conference. Amsterdam, the Netherlands.

*Waters, T.D. & D. Walsh. 2008. Seed Corn Maggot Control in Pacific Northwest, USA, Dry Bulb Onions. Acta Hort. Proceedings of the World Edible Allium Conference. Amsterdam,

Ross, C., H. Ferguson, M. Keller, D. Walsh, K. Weller, and S. Spayd  2007. Influence of the Multicolored Asian Lady Beetle on the Aroma of Concord Grape Juice. J. of Food Qual. 30:855-863.

Brown, J.J., N. Kittelson, E. Hannon, & D. Walsh. 2006. An endemic population of western poplar clearwing moths invades a monoculture of preferred hosts. J. Econ. Entomol. 99: 771-779

Walsh, D.B. & G. Grove. 2005. Repellency and repulsiveness of fungicides, petroleum spray oils, and spray adjuvants to two-spotted spider mites Tetranychus urticae Koch. Plant Health Progress.

Williams, M.M. & D.B. Walsh. 2004 Integrating arthropod herbivory and reduced herbicide use for weed management. Weed Science 52:1018-1025

*Waters, T.D., D.B. Walsh, R.S. Zack, & R.P. Wight. 2004. Ground beetle abundance in disturbed and rehabilitated riparian habitats in Eastern Washington State, USA. Proceedings, International Congress of Entomology. Brisbane Australia.

*Waters, T.D. H. Ferguson, D.B. Walsh. 2003. Ode to a Cricket. Am Entomol.

Zalom, F. G., D. B. Walsh, W. Krueger, and J. Connell. 2002. Tolerance of peach twig borer, Anarsia lineatella, to organophsphate and pyrethroid insecticides. Acta Horticulturae 591:585-590.

Walsh, D.B., D.V. Shaw, F.G. Zalom & K.D. Larson. 2002. Yield reduction caused by two-spotted spider mite (Acari: Tetranychidae) feeding in an advanced-cycle strawberry breeding population. J. Amer. Soc. Hort Sci. 127:230-237.

Walsh, D.B., D.V. Shaw, F.G. Zalom & K.D. Larson. Yield reduction caused by two-spotted spider mite (Acari: Tetranychidae) feeding in an advanced-cycle strawberry breeding population. J. Amer. Hort. Sci. 127(2):230-237.

Zalom, F.G., M. W. Stimann, T.S Arndt, D.B. Walsh, C. Pickel, & W.H. Krueger. 2001. Analysis of permethrin (cis- and trans-isomers) and esfenvalerate on almond twigs and effects of residues on the western orchard predator mite, Galandromus occidentalis Nesbitt. Env. Entomol. 30:70-75

Walsh, D.B. 2000. Potential impacts of riparian habitats on population abundance of pest arthropods. In. P. J. Wigington (ed.) American Water Resources Association. Riparian Ecology and Management in Multi-Land Use Watersheds. pp. 101-106.

Walsh, D.B., F.G. Zalom & D.V. Shaw. 1998. Interaction of the two-spotted spider mite (Acari Tetranychidae) with yield of day-neutral strawberries in California. J. Econ. Entomol. 91:668-675.

Walsh, D.B., F.G. Zalom, N. C. Welch, C. Pickel, & D.V. Shaw. 1997. Pre-transplant cold storage of strawberries: effects on plant vigor, yield and spider mite (Acari: Tetranychidae) Abundance. J. Econ. Entomol. 90(3):818-823.

Walsh, D.B. F. Zalom. & M. Stimmann. 1997. Effects of pyrethroid insecticide residues on almond leaves on the biology of the Western orchard predatory mite Gallendromes occidentalis. II. International Symposium on Pistachios and Almonds. Davis, CA. Acta Horticulturae 470:539-546.

Walsh, D.B., F.G. Zalom, D.V. Shaw, & N. C. Welch. 1996. Effect of strawberry plant physiological status on the translaminar activity of avermectin B1 and its efficacy on the two-spotted spider mite Tetranychus urticae Koch (Acari: Tetranychidae). J, Econ. Entomol. 89(5): 1250-1253.

Pickel, C., F. G. Zalom, D. B. Walsh, & N. C. Welch. 1995. Vacuums provide limited Lygus control in strawberries. Calif. Agric. 49(2):19-22.

Pickel, C., F. G. Zalom, D. B. Walsh, & N. C. Welch. 1994. Efficacy of vacuum machines for Lygus hesperus Knight (Hemiptera: Miridae) control in coastal California strawberries. J.Econ. Entomol.87(6): 1636-1640.

Zalom, F. G., C. Pickel, D. B. Walsh & N. C. Welch. 1993. Sampling for Lygus hesperus Knight (Hemiptera: Miridae) in strawberries. J. Econ. Entomol. 86(4): 1191-1195.

Book Chapters

Walsh, D. B. & F. G. Zalom. 2000. Programma de manejo de plagas de artropodas de las fresa en las regiones productores de California. In. J.Z. Castellenos & F.G. O’Hart (eds.) Fresa, Productivida con Progreso. Zamora, MX. pp.109-122.

Walsh, D.B. 2002. Acaricides. In D. Pimental (ed.). Encyclopedia of Pest Management. Marcel Dekker Press. New York, NY.

Walsh, D. & F. Zalom. 2002. Winter spray oil treatments for control of two-spotted spider mite on strawberries. Spray Oils Beyond 2000. Sydney, Australia.

Zalom, F. & D. Walsh. 2002. Petroleum-derived spray oils: a useful tactic in California IPM systems. Spray Oils Beyond 2000. Sydney, Australia.

Agrichemical and Environmental News, A monthly report on pesticides and related environmental issues.

  • Walsh, D. 2001. Homoptera and Neonicotinyls. Imidacloprid Aids in GWSS Battle in California. Issue No. 186.
  • Walsh, D. 2001. IR-4 Projects 2001 & 2002 (Fungicides). Issue No. 184.
  • Walsh, D. 2001. Dragonflies. Issue No. 184.
  • Walsh, D. 2001. IR-4 Projects 2001 & 2002 (Insecticides and Herbicides). Issue No. 183.
  • Walsh, D. 2001. Thrips and Onions Field Observations 2000. Issue No. 182.
  • Walsh, D. 2001. Lygus bugs. Issue No. 182.
  • Walsh, D., & G. Grove. 2001. Repelled and Repulsed. Two-Spotted Spider Mites React to Agrichemicals. Issue No. 181.
  • Walsh, D. 2001. Thrips. Issue No. 181.
  • Walsh, D., K. Eastwell, D. James, & W. Bentley. 2001. Grape Mealybug. Issue No. 179.
  • Walsh, D. 2000. Riparian Buffer Zones: Summer 2000 Field Study. Issue No. 176.
  • Walsh, D. 2000. IR-4 Priorities Set for 2001. Issue No. 176.
  • Walsh, D. 2000. Pesticide AD-ucation: “Learning” from Madison Ave. Issue No. 175.
  • Walsh, D. 2000. Input Needed Now for 2001 IR-4 Projects. Issue No. 173.
  • Walsh, D. 2000. Changes in WSCPR Funding. Issue No. 173
  • Walsh, D. 2000. The application of moth pheromones. Issue No. 171.
  • Walsh, D. 2000. The biology of moth pheromones. Issue No. 170.
  • Walsh, D. 2000. Noteworthy new products-Insecticides. Issue No. 170.
  • Walsh, D. 2000. Noteworthy new products-Fungicides. Issue No. 169.
  • Walsh, D. 2000. The chemistry of moth pheromones. Issue No. 168.
  • Walsh, D. 2000. Noteworthy new products-Herbicides. Issue No. 168.
  • Walsh D. 2000. Y2K. Its not bugging the bugs. Issue No. 166.
  • Walsh, D. and F. Zalom. 1999. Horticultural Spray Oils: A Useful Tactic for IPM Systems. Issue No. 165.
  • Walsh D. 1999. IR-4: Developing and Delivering Pest Management Solutions for Minor Crop Producers. Issue No. 162.
  • Walsh D. 1999. Implications of buffer zones on agricultural lands; Impacts on Beneficial and Pest Organisms. Issue No. 163.
  • Walsh, D. 1999. IPM An overview. Issue No. 159.
  • Walsh, D. 1999. Codling moth: An IPM model. Issue No. 159.
  • Walsh, D. 1999. Plant sucking mites. Major pests of modern food fiber and ornamental crops. Issue No. 158.
  • Walsh, D. 1999. SFI a boon for Washington State. Issue No. 158.
  • Walsh, D. 1999. Coming soon to a pesticide label near you. Issue No. 158.
  • Walsh, D. 1999. The ecological basis for insecticide resistance. Issue No. 156.
  • Walsh, D. 1999. Spinosad, a lethal brew for pest insects. Issue No. 154.
  • Walsh, D. 1998. What if OPs and carbamates were banned? Issue No. 152.
  • Walsh, D. 1998. Pleased to meet you. Issue No. 150.

Department of Entomology
Washington State University
Irrigated Agriculture Research
and Extension Center

Prosser WA 99350
Phone: 509-786-9287
Fax: 509-786-9370

WSU IPM Website

WSU Prosser Website